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Australopithecus africanus Fossil range: Pliocene
Scientific classification
Kingdom: Animalia Phylum: Chordata Class: Mammalia Order: Primates Family: Hominidae Subfamily: Homininae Genus: Australopithecus Species: A. africanus
Binomial name
†Australopithecus africanus Dart, 1925

Australopithecus africanus was an early hominid, an australopithecine, who lived between 3.3 and 2.4 million years ago in the Pliocene. In common with the older Australopithecus afarensis, A. africanus was slenderly built, or gracile, and was thought to have been a direct ancestor of modern humans. Fossil remains indicate that A. africanus was significantly more like modern humans than A. afarensis, with a more human-like cranium permitting a larger brain and more humanoid facial features.

Main article: Taung Child

Raymond Dart was at Taung near Kimberley, South Africa in 1924 when one of his colleagues spotted a few bone fragments and the cranium on the desk of a lime worker. The skull seemed like an odd ape creature sharing human traits such as eye orbits, teeth, and, most importantly, the hole at the base of the skull over the spinal column (the foramen magnum) indicating a human-like posture. Dart assigned the specimen the name Australopithecus africanus ("southern ape of Africa"). This was the first time the word Australopithecus was assigned to any hominid. Dart claimed that the skull must have been an intermediate species between ape and humans, but his claim about Taung Child was rejected by the scientific community at the time. Sir Arthur Keith suggested that the skull belonged to a young ape, most likely from an infant gorilla.

Main article: Mrs. Ples

Dart's theory was supported by Robert Broom. In 1938 Broom classified an adult endocranial cast having a brain capacity of 485cc, which had been found by G. W. Barlow, as Plesianthropus transvaalensis. On April 17, 1947, Broom and John T. Robinson discovered a skull belonging to a middle-aged female, Sts 5, while blasting at Sterkfontein. Broom classified it also as Plesianthropus transvaalensis, and it was dubbed Mrs. Ples by the press (though the skull is now thought to have belonged to a young male). The lack of facial projection in comparison to apes was noted by Raymond Dart (including from Taung Child), a trait in common with more advanced hominines. Both fossils were later classified as A. africanus.

Like A. afarensis, A. africanus the South African counterpart was generally similar in many traits, a bipedal hominin with arms slightly larger than the legs (a physical trait also found in chimpanzees). Despite its slightly more human-like post cranial features, seen for example in the craniums Mr. Ples and Sts 71, other more primitive features including ape-like curved fingers for tree climbing are also present.

Raymond Dart with Taung skull

Due to other more primitive features visible on A. africanus, some researchers believe the hominin, instead of being a direct ancestor of more modern hominins, evolved into Paranthropus. The one particular robust australopithecine seen as a descendent of A. africanus is Paranthropus robustus. Both P. robustus and A. africanus craniums seem very alike despite the more heavily built features of P. robustus that are adaptations for heavy chewing like a gorilla. A. africanus, on the other hand, had a cranium which quite closely resembled that of a chimp, yet both their brains measure about 400 cc to 500 cc and probably had an ape-like intelligence. A. africanus had a pelvis that was built for slightly better bipedalism than that of A. afarensis. No stone tools of any sort have ever been found in association with australopithecines with the exception of 2.6 million year old Australopithecus garhi.

Charles Darwin suggested that humans had originally evolved from Africa, but during the early 20th century most anthropologists and scientists supported the idea that Asia was the best candidate for human origins. However, the famous Leakey family have argued in favor of the African descent since most hominid discoveries such as the Laetoli footprints were uncovered in Eastern Africa. The species A. africanus with its presumably slightly more Homo-like post cranial features in comparison to A. afarensis is one of several Australopithecine candidates to have evolved into the genus Homo (ie. Homo habilis and Homo rudolfensis by 2.4 million years ago).^{[citation needed]}

• Cranial capacity
• List of human fossils
• List of fossil sites (with link directory)
• List of hominina (hominid) fossils (with images)

• BBC - Dawn of Man (2000) by Robin Mckie| ISBN 0-7894-6262-1
• Early human phylogeny. Smithsonian Institution (2000-2004). Retrieved on April 8, 2007.
• Human ancestry. ArchaeologyInfo.com (1999-2006). Retrieved on April 8, 2007.
• Hilton-Barber, Brett; Berger, Lee R (2004). Field guide to the cradle of humankind, Sterkfontein, Swartkrans, Kromdraai & environs world heritage site, 2nd revised edition. Struik. ISBN 1-77007-065-6. 

• MNSU
• Archaeology Info
• Smithsonian
• Handprint