Boraginaceae

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Boraginaceae
Forget-me-not (Myosotis discolor)
Forget-me-not (Myosotis discolor)
Scientific classification
Kingdom: Plantae
Division: Magnoliophyta
Class: Magnoliopsida
Family: Boraginaceae
Juss.
Genera

many, see text

Boraginaceae Juss. 1789, the Borage or Forget-me-not family, includes a variety of shrubs, trees, and herbs, totaling about 2,000 species in 100 genera found worldwide. A number of familiar plants belong to this family.

The Boraginaceae belong, according to the APG II, among the euasterid I group including the orders Gentianales, Lamiales, and Solanales, but whether they should be assigned to one of these orders or to their own (Boraginales) is still uncertain. Under the older Cronquist system they were included in the Lamiales, but it is now clear that they are no more similar to the other families in this order than they are to families in several other asterid orders. The Boraginaceae are paraphyletic with respect to Hydrophyllaceae and the latter is included in the former in APG II system. In some recent classifications the Boraginaceae are broken up into several families: Boraginaceae s.s., Cordiaceae, Ehretiaceae, Heliotropiaceae, Hydrophyllaceae, and Lennoaceae.

Most though not all members of this family have hairy leaves.

Well-known members include:

  • Actinocarya
  • Adelocaryum
  • Afrotysonia
  • Alkanna
  • Amblynotus
  • Amphibologyne
  • Amsinckia
  • Anchusa
  • Ancistrocarya
  • Anoplocaryum
  • Antiotrema
  • Antiphytum
  • Arnebia
  • Asperugo
  • Auxemma
  • Borago
  • Bothriospermum
  • Bourreria
  • Brachybotrys
  • Brunnera
  • Buglossoides
  • Caccinia
  • Carmona
  • Cerinthe
  • Chionocharis
  • Choriantha
  • Cordia
  • Craniospermum
  • Cryptantha
  • Cynoglossopsis
  • Cynoglossum
  • Cynoglottis
  • Cysostemon
  • Dasynotus
  • Decalepidanthus
  • Echiochilon
  • Echiostachys
  • Echium
  • Ehretia
  • Elizaldia
  • Embadium
  • Eritrichium
  • Gastrocotyle
  • Gyrocaryum
  • Hackelia
  • Halacsya
  • Halgania
  • Heliocarya
  • Heliotropium
  • Heterocaryum
  • Huynhia
  • Ivanjohnstonia
  • Ixorhea
  • Lacaitaea
  • Lappula
  • Lasiarrhenum
  • Lasiocaryum
  • Lepechiniella
  • Lepidocordia
  • Lindelophia
  • Lithodora
  • Lithospermum
  • Lobostemon
  • Macromeria
  • Maharanga
  • Mairetis
  • Mattiastrum
  • Mertensia
  • Metaeritrichium
  • Microcaryum
  • Microula
  • Mimophytum
  • Moltkia
  • Moltkiopsis
  • Moritzia
  • Myosotidium
  • Myosotis
  • Neatostema
  • Nesocaryum
  • Nogalia
  • Nomosa
  • Nonea
  • Ogastemma
  • Omphalodes
  • Omphalolappula
  • Omphalotrigonotis
  • Onosma
  • Onosmodium
  • Oxyosmyles
  • Paracaryum
  • Pardoglossum
  • Patagonula
  • Pectocarya
  • Pentaglottis
  • Perittostema
  • Phacelia
  • Plagiobothrys
  • Pseudomertensia
  • Psilolaemus
  • Pteleocarpa
  • Pulmonaria
  • Rindera
  • Rochefortia
  • Rochelia
  • Rotula
  • Saccellium
  • Scapicephalus
  • Selkirkia
  • Sericostoma
  • Sinojohnstonia
  • Solenanthus
  • Stenosolenium
  • Stephanocaryum
  • Suchtelenia
  • Symphytum
  • Thaumatocaryum
  • Thyrocarpus
  • Tianschaniella
  • Tiquilia
  • Tournefortia
  • Trachelanthus
  • Trachystemon
  • Trichodesma
  • Trigonocaryum
  • Trigonotis
  • Ulugbekia
  • Valentiniella
Wikimedia Commons has media related to:

  • Diane, N., H. Förther, and H. H. Hilger. 2002. A systematic analysis of Heliotropium, Tournefortia, and allied taxa of the Heliotropiaceae (Boraginales) based on ITS1 sequences and morphological data. American Journal of Botany 89: 287-295 (online abstract here).
  • Gottschling, M., H. H. Hilger 1, M. Wolf 2, N. Diane. 2001. Secondary Structure of the ITS1 Transcript and its Application in a Reconstruction of the Phylogeny of Boraginales. Plant Biology (Stuttgart) 3: 629-636.

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