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Crocodylomorpha Fossil range: Late Triassic - Recent
Protosuchus, an early crocodylomorph
Scientific classification
Kingdom: Animalia Phylum: Chordata Class: Sauropsida Infraclass: Archosauromorpha (unranked) Crurotarsi Superorder: Crocodylomorpha Hay, 1930
Orders
see text

The Crocodylomorpha are an important group of archosaurs that include the crocodilians and their extinct relatives.

When their extinct species and stem group are examined, the crocodylian lineage (clade Crurotarsi) proves to have been a very diverse and adaptive group of reptiles. Not only are an ancient group of animals, at least as old as the dinosaurs, they also evolved into a great variety of forms. The earliest forms, the Sphenosuchians, evolved during the Late Triassic, and were highly gracile terrestrial forms built like greyhounds. Several terrestrial species during the Cretaceous evolved herbivory, such as Simosuchus clarki and Chimaerasuchus paradoxus. During the Jurassic and the Cretaceous marine forms in the family Metriorhynchidae such as Metriorhynchus evolved forelimbs that were paddle-like and had a tail similar to modern fish. Dakosaurus andiniensis a closely related species to Metriorhynchus had a skull that was adapted to eat large marine reptiles.

During Mesozoic and early Tertiary times the Crocodylomorpha were far more diverse than they are now. Triassic forms were small, lightly built, active terrestrial animals. These were supplanted during the early Jurassic by various aquatic and marine forms. The Later Jurassic, Cretaceous, and Tertiary saw a wide diversity of terrestrial and semi-aquatic lineages. "Modern" crocodilians do not appear until the Late Cretaceous.

The Crocodylomorpha are defined phylogenetically by Sereno 2005 as "The most inclusive clade containing Crocodylus niloticus (Laurenti 1768) but not Poposaurus gracilis Mehl 1915, Gracilisuchus stipanicicorum Romer 1972, Prestosuchus chiniquensis Huene 1942, Aetosaurus ferratus Fraas 1877."

This a stem-based definition and therefore includes all taxa closer to extant crocodilians than to other crurotarsan clades.

Historically, all known living and extinct crocodiles were indiscriminately lumped into the order Crocodilia. But it is now known that this is erroneous, based on the uniqueness of the crocodilian morphology. Thus, the order Crocodilia is nowadays restricted to the living species and close extinct relatives such as Mekosuchus.

The old Crocodilia was subdivided into the suborders:

• Eusuchia (which includes crown-group Crocodylia)
• Mesosuchia ("primitive crocodiles")
• Protosuchia ("proto-crocodiles")
• Thalattosuchia ("sea crocodiles")

Mesosuchia is a paraphyletic group as it does not include eusuchians (which nest within Mesosuchia). Mesoeucrocodylia was the name given to the clade that contains mesosuchians and eusuchians (Whetstone and Whybrow, 1983).

Mikko's Phylogeny Archive provided this detailed cladogram, after Clark (1994), Ortega et al. (2000), and Turner & Calvo (2005):

Crocodyliformes
  |--Eopneumatosuchus
  |-?Microchampsa
  `-+-?Artzosuchus
    |--Protosuchia
    `--+--Shartegosuchidae
       `--Mesoeucrocodylia
           |-?Hsisosuchus
           `--+-?Gobiosuchidae
              `--+-?Simosuchus
                 |--Ziphosuchia
                 `--+-?Uruguaysuchus
                    `--+--Araripesuchus
                       `--+--Peirosauridae
                          `--+--Trematochampsidae
                             `--Neosuchia
                                 |-?Elosuchidae
                                 |--Peirosauridae
                                 `--+--Comahuesuchus
                                    |--Itasuchus
                                    |--Atoposauridae
                                    `--+--+--Eutretauranosuchus
                                       |  `--+-?+--Dyrosauridae
                                       |     |  `--+--Sarcosuchus
                                       |     |     `--Terminonaris
                                       |     `--+--Pholidosauridae
                                       |        `--+--Thalattosuchia
                                       |           `--+--Pelagosuchus
                                       |              `--+--Teleosauridae
                                       |                 `--Metriorhynchidae
                                       `--+--Goniopholididae
                                          `--+--Bernissartia
                                             `--+--Gilchristosuchus
                                                |--Rugosuchus
                                                |-?Paralligatoridae
                                                `--Eusuchia
                                                    |--Hylaeochampsa
                                                    `--+--Allodaposuchus
                                                       `--Crocodilia

The previous definitions of Crocodilia and Eusuchia do not accurately resemble the evolution of the group. The only order-level taxon that is currently considered valid is Crocodilia in the present definition. Prehistoric crocodiles are represented by many taxa, but since few major groups of the ancient forms are recognizable, a decision where to delimit new order-level clades is not yet possible. (Benson & Clark, 1988).

• Benton, M. J. (2004), Vertebrate Palaeontology, 3rd ed. Blackwell Science Ltd
• Hay, O. P. 1930 (1929-1930). Second Bibliography and Catalogue of the Fossil Vertebrata of North America. Carnegie Institution Publications, Washington, 1,990 pp.
• Sereno, P. C. 2005. Stem Archosauria—TaxonSearch [version 1.0, 2005 November 7]
  • Taxon Search - Crocodylomorpha

• Crocodylomorpha - webpages by Ross Elgin on the University of Bristol server
  • Major subgroups classification (used here)
• Crocodylomorpha from Palaeos
  • Technical definition
• Crocodylomorpha - hyperlinked cladogram at Mikko's Phylogeny Archive

v • d • e Archosauromorphs
Primitive Archosauromorphs	Euparkeriidae • Erythrosuchidae • Proterochampsidae • Proterosuchidae • Choristodera • Prolacertiformes • Rhynchosauria • Trilophosauria
Crurotarsi Archosaurs	Ornithosuchidae • Aetosauria • Phytosauria • Rauisuchia • Crocodylomorpha • Crocodilia
Avemetatarsalia and Ornithodira Archosaurs	Scleromochlus • Pterosauria • Dinosauromorpha • Dinosauria • Ornithischia • Saurischia • Aves
Avian Archosaurs	Avialae • Archaeopteryx • Confuciusornis • Ichthyornis • Enantiornithes • Hesperornithes • Neornithes • Paleognathae • Neognathae