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Hummingbird
A variety of hummingbirds from Ernst Haeckel's 1904 Kunstformen der Natur (Artforms of Nature)
Scientific classification
Kingdom: Animalia Phylum: Chordata Class: Aves Order: Apodiformes Family: Trochilidae Vigors, 1825
Subfamilies
Phaethornithinae Trochilinae For a taxonomic list of genera, see: List of hummingbirds in taxonomic order For an alphabetic species list, see: Alphabetic species list

Hummingbirds are small birds in the family Trochilidae, native only to the Americas. They are known for their ability to hover in mid-air by rapidly flapping their wings, 15–80 times per second (depending on the species). Capable of sustained hovering, the hummingbird has the ability to fly deliberately backwards (this is the only group of birds able to do so^[1]) or vertically, and to maintain position while drinking nectar or eating tiny arthropods from flower blossoms. They are named for the characteristic hum made by their wings.

A hummingbird with its tongue out.

Hummingbirds are small birds with a long, thin bills. This elongated bill is one of the defining characteristics of the hummingbird, which, with an extendable, bifurcated tongue, has evolved in order to allow the bird to feed upon nectar deep within flowers. A hummingbird's lower bill also has the ability to flex downward to create a wider opening, facilitating the capture of insects in the mouth rather than at the tip of the beak.^[2]

Iridescent plumage in a male Anna's Hummingbird.

The bee Hummingbird (Mellisuga helenae) is the smallest bird in the world, weighing 1.8 grams (0.06 ounces) and measuring about 5 cm (2 inches). A more typical hummingbird, such as the Rufous Hummingbird (Selasphorus rufus), weighs approximately 3 g (0.106 ounces) and has a length of 10–12 cm (3.5–4 inches). The largest hummingbird is the Giant Hummingbird (Patagona gigas), with some individuals weighing as much as 24 grams (0.85 ounces) and measuring 21.5 cm (8.5 inches).

They display sexual dimorphism, as male hummingbirds are usually more brightly colored, while females of most species display more cryptic coloration.^[3] Most males have iridescent plumage, in metallic red, orange, green and/or blue.

Hummingbird at a flower

Hummingbirds are attracted to many flowering plants — shrimp plants, bee balm, Heliconia, Buddleja, Hibiscus, bromeliads, cannas, verbenas, honeysuckles, salvias, pentas, fuchsias, many penstemons, and others. It is often stated that they are especially attracted to red and yellow flowers. Once attracted to a garden, hummingbirds may find flowers of other colors more attractive. The location and growing season should determine choices of the plants selected for a garden to attract hummingbirds. They feed on the nectar of these plants and are important pollinators, especially of deep-throated flowers. Hummingbird also take insects or spiders, especially when feeding young.

Hummingbirds are specialized nectarivores (Stiles, 1981) and are tied to the ornithophilous flowers they feed upon. They are often very coevolved with a small number of flower species. Hummingbird at a flower of Callistemon citrinus

A trail of wake vorticies generated by a hummingbird's flight. Discovered after training a bird to fly through a cloud of neutrally buoyant helium-filled soap bubbles and recording airflows in the wake with stereo photography.^[4]

Female Black-chinned Hummingbird

Hummingbirds fly particularly well in wind tunnels and in front of cameras. For this reason, hummingbird flight has been studied intensively from an aerodynamic perspective with high-speed video cameras.

Writing in Nature, the biomechanist Douglas Warrick and coworkers studied the Rufous Hummingbird, Selasphorus rufus, in a wind tunnel using particle image velocimetry techniques and investigated the lift generated on the bird's upstroke and downstroke.

They concluded that their subjects produced 75% of their weight support during the down-stroke and 25% during the up-stroke: many earlier studies had assumed (implicitly or explicitly) that lift was generated equally during the two phases of the wingbeat cycle, as is the case of insects of a similar size. This finding shows that hummingbirds' hovering is similar to, but distinct from, that of hovering insects such as the hawk moths. ^[5]

The Giant Hummingbird's wings beat 8–10 beats per second, the wings of medium-sized hummingbirds beat about 20–25 beats per second and the smallest beat 70 beats per second.

Anna's Hummingbird ,Calypte anna performs Personal grooming

With the exception of insects, hummingbirds while in flight have the highest metabolism of all animals, a necessity in order to support the rapid beating of their wings. Their heart rate can reach as high as 1,260 beats per minute, a rate once measured in a Blue-throated Hummingbird [1]. They also typically consume more than their own weight in food each day, and to do so they must visit hundreds of flowers daily. At any given moment, they are only hours away from starving.

However, they are capable of slowing down their metabolism at night, or any other time food is not readily available. They enter a hibernation-like state known as torpor. During torpor, the heart rate and rate of breathing are both slowed dramatically (the heart rate to roughly 50–180 beats per minute), reducing their need for food. Most organisms with very rapid metabolism have short lifespans, however hummingbirds have been known to survive in captivity for as long as 17 years.

Studies of hummingbirds' metabolism are highly relevant to the question of whether a migrating Ruby-throated Hummingbird can cross 800 km (500 miles) of the Gulf of Mexico on a nonstop flight, as field observations suggest it does. This hummingbird, like other birds preparing to migrate, stores up fat to serve as fuel, thereby augmenting its weight by as much as 40–50 percent and hence increasing the bird's potential flying time.^[6]

Hummingbirds are found only in the Americas, from southern Alaska and Canada to Tierra del Fuego, including the Caribbean. The majority of species occur in tropical Central and South America, but several species also breed in temperate areas. Excluding vagrants, sometimes from Cuba or the Bahamas, only the migratory Ruby-throated Hummingbird breeds in eastern North America. The Black-chinned Hummingbird, its close relative and another migrant, is the most widespread and common species in the western United States and Canada.

Most hummingbirds of the U.S. and Canada migrate to warmer climates in the northern winter, but some remain in the warmest coastal regions. Some southern South American forms also move to the tropics in the southern winter.

The Rufous Hummingbird shows an increasing trend to migrate east to winter in the eastern United States, rather than south to Central America, as a result of increasing survival prospects provided by artificial feeders in gardens. In the past, individuals that migrated east would usually die, but now many survive, and their changed migration direction is inherited by their offspring. Provided sufficient food and shelter is available, they are surprisingly hardy, able to tolerate temperatures down to at least -4 °C (25 °F).

Hummingbird nest with two chicks in Santa Monica, California

Calliope Hummingbird feeding two chicks in Grand Teton National Park

As far as is known, male hummingbirds do not take part in nesting. Most species make a neatly woven cup in a tree branch. Two white eggs are laid, which despite being the smallest of all bird eggs, are in fact large relative to the hummingbird's adult size. Incubation is typically 14–19 days. The nest is usually about the size of a pocket watch.

Hummingbird feather

Traditionally, hummingbirds are placed in the order Apodiformes, which also contains the swifts. Hummingbirds' wings are hollow and fragile, making fossilization difficult and leaving their evolutionary history a mystery. Some scientists also believe that the hummingbird evolved relatively recently. Scientists also theorize that hummingbirds originated in South America, where there is the greatest species diversity. Brazil and Ecuador contain over half of the known species.

There are between 325 and 340 species of hummingbird, depending on taxonomic viewpoint, historically divided into two subfamilies, the hermits (subfamily Phaethornithinae, 34 species in six genera), and the typical hummingbirds (subfamily Trochilinae, all the others). However, recent phylogenetic analyses by Mcguire et al (2007) and colleagues suggest that this division is slightly inaccurate, and that there are nine major clades of hummingbirds: the Topazes, the hermits, the Mangoes,the coquettes, the Brilliants, the Giant Hummingbird (Patagonia gigas), the Mountain Gems, the Bees, and the Emeralds. The Topazes (Topaza pella and Florisuga mellivora) have the oldest split with the rest of the hummingbirds.

The modern diversity of hummingbirds is thought by evolutionary biologists to have evolved in South America, as the great majority of the species are found there. However, the ancestor of extant hummingbirds may have lived in parts of Europe to what is southern Russia today.

Genetic analysis^{[citation needed]} has indicated that the hummingbird lineage diverged from their closest relatives some 35 million years ago, in the Late Eocene, but fossil evidence is limited. Fossil hummingbirds are known from the Pleistocene of Brazil and the Bahamas—though neither has yet been scientifically described—and there are fossils and subfossils of a few extant species known, but until recently, older fossils had not been securely identifiable as hummingbirds.

In 2004, Dr. Gerald Mayr of the Senckenberg Museum in Frankfurt am Main identified two 30-million-year-old hummingbird fossils and published his results in Nature.^[7] The fossils of this primitive hummingbird species, named Eurotrochilus inexpectatus ("unexpected European hummingbird"), had been sitting in a museum drawer in Stuttgart; they had been unearthed in a clay pit at Wiesloch-Frauenweiler, south of Heidelberg, Germany and, because it was assumed that hummingbirds never occurred outside the Americas, were not recognized to be hummingbirds until Mayr took a closer look at them.

Fossils of birds not clearly assignable to either hummingbirds or a related, extinct family, the Jungornithidae, have been found at the Messel pit and in the Caucasus, dating from 40–35 mya, indicating that the split between these two lineages indeed occurred at that date. The areas where these early fossils have been found had a climate quite similar to the northern Caribbean or southernmost China during that time. The biggest remaining mystery at the present time is what happened to hummingbirds in the roughly 25 million years between the primitive Eurotrochilus and the modern fossils. The astounding morphological adaptations, the decrease in size, and the dispersal to the Americas and extinction in Eurasia all occurred during this timespan. DNA-DNA hybridization results ^[8] suggest that the main radiation of South American hummingbirds at least partly took place in the Miocene, some 12–13 mya, during the uplifting of the northern Andes.

• Alphabetical list, sortable by common or binomial name
• List of hummingbirds in taxonomic order

Hummingbirds sometimes fly into garages and become trapped. It is widely believed that this is because they mistake the hanging (usually red-color) door-release handle for a flower, although hummingbirds can also get trapped in enclosures that do not contain anything red. Once inside, they may be unable to escape because their natural instinct when threatened or trapped is to fly upward. This is a life-threatening situation for hummingbirds, as they can become exhausted and die in a relatively short period of time, possibly as little as an hour. If a trapped hummingbird is within reach, it can often be caught gently and released outdoors. Alternatively, if offered a perch (such as a broom or a long branch) by a person that does not make any sudden movements, the hummingbird may land on the perch. This allows the person to slowly, calmly carry the bird outside to fly away safely.

The diet of hummingbirds requires an energy source (typically nectar) and a protein source (typically small insects). Providing many plants that carry blooms used by hummingbirds is the safest way to provide nectar for hummingbirds. Through careful plant selection, gardens may contain plants that bloom at different times to attract hummingbirds throughout the seasons they are present in an area. Placing these plants near windows affords a good view of the birds. Hummingbirds will also take synthetic nectar from artificial feeders. Such feeders allow people to observe and enjoy hummingbirds up-close while providing the hummingbirds with a reliable supply of nectar, especially when flower blossoms are less abundant. Maintaining cleanliness of the feeder is essential for the health of the birds. Homemade nectar can be made from 1 part white, granulated table sugar to 4 parts water, boiled to make it easier to dissolve the sugar and to purify the solution so that it will stay fresh longer. The cooled nectar is then poured into the feeder.

Hummingbirds will either hover or perch to feed, and red feeders are preferred.

Hummingbird hovering to feed at a red feeder with yellow "flowers"

Things to avoid using in feeders include honey, which should not be used because it is prone to culture bacteria dangerous to hummingbirds.^[9] Artificial sweeteners should also be avoided because, although the hummingbirds will drink it, they will be starved of the calories they need to sustain their metabolism. Some commercial hummingbird foods contain red dyes and preservatives, which are unnecessary and possibly dangerous to the birds, so dyes and preservatives should be avoided because neither have been studied for long-term effects on hummingbirds. While it is true that bright colors, especially red, initially attract hummingbirds more quickly than others, it is better to use a feeder that has some red on it, rather than coloring the liquid offered in it. It is possible that red dye is harmful to hummingbirds.^[10] Commercial nectar mixes may contain small amounts of mineral nutrients which are useful to hummingbirds, but hummingbirds get all the nutrients they need from the insects they eat, not from nectar, so the added nutrients also are unnecessary. Authorities on hummingbirds recommend that if you use a feeder, use just plain sugar and water.^[11]

A hummingbird feeder should be easy to refill and keep clean. Prepared nectar can be refrigerated for 1–2 weeks before being used, but once placed outdoors it will only remain fresh for 2–4 days in hot weather, or 4–6 days in moderate weather, before turning cloudy or developing mold. If the feeder is in a shady area the nectar will last longer without spoiling. When changing the nectar, the feeder should be rinsed thoroughly with warm tap water, flushing the reservoir and ports to remove any contamination or sugar build-up. If dish soap is used, it always needs extra rinsing so that no residue is left behind. The feeder can be soaked in dilute chlorine bleach if black specks of mold appear and rinsed with clear water.

Other animals are also attracted to hummingbird feeders. It is a good idea to get a feeder that has very narrow ports, or ports with mesh-like "wasp guards", to prevent bees and wasps from getting inside where they get trapped. Orioles, woodpeckers, banaquits, and other animals are known to drink from hummingbird feeders, sometimes tipping them and draining the liquid. If this becomes a problem, it is possible to buy feeders which are specifically designed to support their extra weight and which hummingbirds will use too. If ants find your hummingbird feeder, one solution is to install an "ant moat", which is available at specialty garden stores and online, or tanglefoot can be used to trap the ants, provided it is applied in a location totally inaccessible to the hummingbirds. You can also place vaseline on the pole that holds the feeder to trap ants on the path that they create.

Sometimes a large hummingbird drives its smaller brethren away from a feeder. An effective solution is to put out a second feeder that contains a slightly lower sugar concentration. Hummingbirds can detect a feeding source that is denser in sugar by only a few percent, and the more aggressive bird will make that feeder its own. The smaller birds will flock to the remaining feeder.

• The Aztec god Huitzilopochtli is often depicted as a hummingbird. The Nahuatl word huitzil (hummingbird) is an onomatopoeic word derived from the sounds of the hummingbird's wing-beats and zooming flight.

Aerial photograph of hummingbird image at Nazca in Peru

• One of the Nazca Lines, displayed at right, depicts a hummingbird.
• The Ohlone tells the story of how a Hummingbird brought fire to the world. See article at the National Parks Conservation Association's website for a recounting.
• Trinidad and Tobago is known as "The land of the hummingbird," and a hummingbird can be seen on that nation's coat of arms,1 cent coin as well as its national airline, "Caribbean Airlines".
• Many popular songs have been written under the title "Hummingbird", including separate works by B.B. King, Wilco, Leon Russell, John Mayer, Frankie Laine, Cat Stevens, Seals and Crofts, Merzbow and Yuki.
• Several companies use a hummingbird as a corporate logo. Such companies include Canadian CHC Helicopter Corporation, British Colibri, and American Hummingbird Scientific.
• In Brazil a black hummingbird of any kind, especially when found inside the house, is a sign of a death in the family.

• Hummingbird Hawk-moth (Macroglossum stellatarum)
• Hemaris, another genus of sphinx moths confused with hummingbirds
• Bird feeder, for information about hummingbird feeders.

• del Hoyo, J.; Elliott, A. & Sargatal, J. (editors) (1999): Handbook of Birds of the World, Volume 5: Barn-owls to Hummingbirds. Lynx Edicions, Barcelona. ISBN 84-87334-25-3

• Gerwin, John A. & Zink, Robert M. (1998): Phylogenetic patterns in the Trochilidae. Auk 115(1): 105-118. fulltextPDF (936 KiB)

McGuire, J. A., Witt, C. C., Altshuler, D. L., and Remsen Jr., J. V. 2007. Phylogenetic systematics and biogography of hummingbirds: Bayesian and maximum likelihood analyses of partitioned data and selection of an appropriate partitioning strategy. Systematic Biology, 56: 837-856.

• Meyer de Schauensee, Rodolphe (1970): A Guide to Birds of South America. Livingston, Wynnewood, PA.

• Stiles, Gary. 1981. Geographical Aspects of Bird Flower Coevolution, with Particular Reference to Central America. Annals of the Missouri Botanical Garden 68:323-351.

Magnificent Hummingbird—Guadalupe, Panama	A male Costa's Hummingbird, showing its plumage to good effect	Male Green Violet-ear	A hovering Rufous Hummingbird on Saltspring Island
The size of a hummingbird	hummingbird among flowers	hummingbird among flowers	hummingbird among flowers
two males fighting	Hummingbird among Crocosmia		Grooming

Wikimedia Commons has media related to:

Trochilidae

• Hummingbirds.net
• The Hummingbird Society
• Ruby Throat Hummingbird Project
• Ecuador Hummingbirds and Climate Change
• Hummingbird species
• The Hummingbirds Video Guide