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Vespidae
Southern yellowjacket (Vespula squamosa) queen
Scientific classification
Kingdom: Animalia Phylum: Arthropoda Class: Insecta Order: Hymenoptera Suborder: Apocrita Superfamily: Vespoidea Family: Vespidae
Subfamilies
Eumeninae: potter wasps Euparagiinae Masarinae: pollen wasps Polistinae: paper wasps Stenogastrinae Vespinae: yellowjackets, hornets

The Vespidae are a large (nearly 5,000 species), diverse, cosmopolitan family of wasps, including nearly all the known eusocial wasps and many solitary wasps. Each social wasp colony includes a queen and a number of female workers with varying degrees of sterility relative to the queen. In temperate social species, colonies usually only last one year, dying at the onset of winter. New queens and males (drones) are produced towards the end of the summer, and after mating, the queens hibernate over winter in cracks or other sheltered locations. The nests of most species are constructed out of mud, but polistines and vespines use plant fibers, chewed to form a sort of paper (also true of some stenogastrines).

The subfamilies Polistinae and Vespinae are composed solely of eusocial species, while Eumeninae, Euparagiinae, and Masarinae are all solitary; Stenogastrinae contains a variety of forms from solitary to social.

In Polistinae and Vespinae, rather than consuming prey directly, prey are masticated and fed to the larvae, and the larvae, in return, produce a clear liquid (with high amino acid content) which the adults consume; the exact amino acid composition varies considerable among species, but it is considered to contribute substantially to adult nutrition^[1].

The nest pictured below was made by either the bald-faced hornet (American), Dolichovespula maculata (sometimes classified as Vespula maculata), or the aerial yellowjacket (that is aerial-nesting yellowjacket), Dolichovespula arenaria. The wasp pictured at right is a yellowjacket queen, Vespula squamosa.

European wasp	A hornet nest	A different angle	Polistes nest
Another angle	Another angle	Vespa tropica from India	Vespa crabro (European hornet) stripping wood from a fence for use in nest construction

• Evolutionary history of social behavior in Vespids
• Vespidae of the world - Iconographic work

1. ^ Hunt, J. H., I. Baker, and H. G. Baker. 1982. Similarity of amino acids in nectar and larval saliva: the nutritional basis for trophallaxis in social wasps. Evolution 36: 1318-1322

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